36 “omnivores,” spanning the second, levels. third and fourth trophic Detritus was shown as providing direct linkage only between the phytoplankton of the pelagic and the algae of the benthic divisions. Linkage between the carnivorous forms of each division waa depicted as through the "omnivore" classification or via the fifth trophic level, the transient carnivores. In a study of the bottom sediments of Rongelap Atoll lagoon, Anikouchine (1961) concluded that the sediments are derived almost entirely from organisms living within the lagoon. There was no significant loss of sediments from the lagoon nor exchange of sedimentary material between the lagoon and oceanic environments. In such a system, insoluble atmos- pheric particles deposited in the lagoon and having sufficient density to settle through the water would be expected to be retained in the lagoon. Finely dispersed, suspended particles would be subject to exchange with the oceanic environment along with the water in which they were suspended. However, particles in these smaller size ranges are also subject to surface phenomena such as flocculation and adsorption, and consequently to gravitational and biological influences tending to retain them within the lagoon.

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