17 Table 8 ABO, MN,Rh-Hr, and Duffy-Kell-Diego Frequency Among Marshallese and Polynesians ABO system — Marshallese (present study) Marshallese (Simmons’*) Polynesians (Simmons and Graydon*') Group Gene frequency No. O A B 129 581 186 15 678 $2.2 21.4 122.1 138 39.1 60.9 0 Rh, Rh, Rhrh Rh, 97.7 23 0 Marshallese (Simmons?) 90.6 0.7 0.3 Polvnesians (Simmons and Graydon’') 19.6 0.7 29.7 (present study) 4) Duffy System: In this system 89.2% Duffy (Fy*) positive bloods were found. A previous report of 100% Duffy (Fy*) Positive reactions’® (in 30 specimens that had been stored for 16 months) indicates a need for verifica- tion and clarification. 5) Other Systems: Kell tests were 100% negative as previously reported. Diego* tests were 100% negative. The failure to demonstrate the Diego factor in anv of the studies conductedin this area ofthe world is noteworthy. To date its absence in Polynesians,?' Maoris,** and now in Marshallese becomes a significant finding in view ofits occur- rence in Mongoloids, Eskimos, and Amerin- dians,**-** to whom Heyerdahl**credits the popu- lation of the Polynesian Islands. The gene frequency comparisons with other reports from this area are shown in Table 8. The above findings indicate a rather homogenous population of the Marshall Islands with ex- *We are indebted to Dr. Philip Levine who supplied the anu Dit serum and Dr. Miguel Layrisse who supplied the Di* positive ceils for control. Type Genefrequency Pe Fe r. M MN N m n 8.2 0.789 0.116 0095 62 155 783 0.14 0.86 5.3 0.723 0.135 0.134 (10) (19) (71) 0.22 0.78 0 0.626 0.374 0.10 47.8 32.6 0435 0.565 Phenotype Rh-Hr Marshallese MNsystem 19.6 Genefrequency Rh,Rh, R' R? Duffy, Kell, Diego R° Fy+ K+ 0.985 0 0.15 89.2 0 0 8.0 0.951 0.04 0.006 - - 50.0 0.449 0.543 0.007 0 0 100 74.6 D+ tremes of gene frequencies. With some reservations because of the relatively small samplings, the following facts are of interest in the blood groupings of the Marshallese. 1) The extremely high frequency of the O gene (78.9%). 2) The extremely low frequency of the M gene (14%). 3) The highest incidence of the R' chromosome yet reported (98.5%). 4) The presence of 10.8% of Duffy (Fy*) negatives. 5) The absolute absence of Kell and Diego factors. 6) A single example of A,B in this area. The investigations of numerous authors, compiled and summarized by Mourant,'* most nearly relate these blood groupings to those found in Southeast Asia and Indonesia, whererelatively frequent B genes are found, a high N frequency exists, and a similar high frequency of the R' chromosomeis seen. The absolute absence of the Diego factor, the extremely low incidence of the M gene, andthe © unusually high R' chromosomefrequencyof the

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